Many natural ecosystems are periodically exposed to fire.  After a fire, there is often reduced competition and increased nutrient availability (from ash etc.).  The plants and flowers that grow after a fire are visited more often by pollinators, such as bees and other insects.  This can result in increased production of fruits and seeds. Bushfires have been part of certain australian ecosystems for thousands of years and some native species are ‘fire adapted’.  They have come to 'rely' on fires as a means of reproduction and / or  dispersal. Whilst no one fire can be attributed to climate change alone, rising temperatures and aridity, lengthening of the ‘fire season’, combined with bursts of extreme ‘fire weather’, all combine to suggest climate change is implicated. As the frequency of fires increases, the possible benefits of fire to such ecosystems / species are being lost. Fire can help with the physical dispersal of seeds from the parent plant.  In some parts of the world, such as South Africa and Australia, fire and / or smoke can be the stimulus for seed dispersal and subsequent germination.  Plants such as some species of Protea, Banksia, certain members of the myrtle family (e.g. some Eucalypts), and some Pines and Sequoias 'make use' of fire to disperse their seeds. Seed dispersal involving fire is termed serotiny.  Many of these plants produce woody fruits or cones in which the seeds are held.  The mechanism underlying seed release varies but can be due to a resin that ‘seals’ the seeds inside the fruit or cone.  The resin ‘melts’ / liquefies on exposure to heat releasing the seed or there may be a structure called a seed separator (as in Banksia).  Serotinous conifers (like lodgepole pine), have mature cones in which the cone scales are naturally sealed shut with resin.   Most of the seeds stay in the canopy until the cones reach 122-140o F  (i.e 50 to 60oC).  At these temperatures, heat / fire  melts the resin and  the cone scales open to expose the seed. The seed can then drop or drift to a burned but cooling ash-rich soil bed. The seeds do well on the burnt soil available to them as the site offers reduced competition, more light, warmth plus the nutrients from the burning of leaves and litter.  Some species align their germination to immediate post-fire conditions - stimulated by chemicals present in the smoke.  The organic compounds karrikins,  products of the degradation of cellulose are  a germination ‘cue’ for some species.  Karrikins are thought to be present on the soil surface after a fire.  When it rains,  the karrikins are 'washed' into the soil, and seeds present in the soil seed bank are then stimulated to germinate. Thanks to Steve Sangster and John Cameron for images of woodland fire.  

Many plants have a distinctive scent, think of sweet peas, jasmine or honeysuckle, or stand next to a pine tree on a warm, summer’s day.  The scent is due to the release of volatile organic compounds (VOC’s, often oils), produced by specific tissues or glands.  Often it is the nectaries within flowers that produce the scent, apart from their ‘job’ of producing the sugary nectar.  The nectaries may be found on almost any structure within a flower - petals, sepals, stamens, ovary*. The location of nectaries varies from species to species.  There are other structures that can produce scent, for example, trichomes, and osmophores. Osmophores are clusters of cells specialising in scent production.  Any part of a plant can release scent, for example, the leaves of eucalyptus, lavender or myrtle. The scent of a plant may include a variety of VOC’s, indeed there may be dozens of different organic compounds contributing to a particular scent.  Many of these compounds are terpenoids (isoprenoids).  They contribute to the scent of eucalyptus oil, lavender oil and the flavours of cinnamon and ginger. Scent may have a number of functions.  It may be released to attract specific pollinators - moths, butterflies, bees, hoverflies etc. (who have learned to recognise the scent).  The production of VOCs can be modulated, for example,  scent production may be turned off when a flower is pollinated.  A scent may also unfortunately be a signal to herbivorous insects to ‘come and feed’. So, scent have positive or negative effects. A scent may be produced to deter herbivory by certain insects.  Sometimes, plants have a different approach. For example,  when pollen beetles feed on oil seed rape, the rapeseed releases VOCs which attract the attention of other insects.  Specifically, those that will lay their eggs in the larvae of the pollen beetles. These insects are usually from the same family as bees, wasps and ants - the Hymenoptera (insects with membranous wings and a ‘narrow waist’).   The pollen beetle larvae are then ‘eaten’ from the inside by the developing parasitoid larva.  The release of VOC’s is affected by a number of factors temperature, light, circadian rhythms, physical damage and drought.  As the temperature increases so the amount of VOCs released increases (usually). This may be experienced in coniferous woodland.  Conifers give off a variety of volatile oils (i.e. biogenic VOC’s) that contribute to a unique aroma and the formation of aerosols found in the air in and around such woodlands and forests; it is most noticeable in warm weather.  [An aerosol is a ‘mixture’ of very small particles (solid or liquid) in air; other examples of aerosols include mist, cigarette smoke, or car exhaust fumes]. In snapdragons, the most scent is emitted at noon which tends to coincide with pollinator activity, in contrast tobacco plants scent release is in the evening / night when hawkmoth are active.  Drought reduces the ability of plants (like rosemary and thyme) to produce / release VOC’s, this in turn, has been observed to affect which pollinators visit their flowers.  Nectaries located within the flowers of a plant are sometimes referred to as nuptial nectaries, whereas those found in other parts are termed extra-nuptial.  

We know that forests are important to all life on the planet.  They have often been referred to as the ‘lungs of the earth’, a reference to the fact that they produce vast quantities of oxygen - which is essential for respiration for so many forms of life.  They also take up carbon dioxide and ‘fix’ it into complex organic molecules - from starches, to cellulose and lignin.  Thus, the carbon is locked away for months, years or even millennia.  The equatorial forests of Brazil and Sumatra are species rich, incredibly diverse, but deforestation and the expansion of agriculture are threats to many biodiverse, forested areas across the world. As so many forests and woodlands have been felled, there is now a movement to plant millions and millions of trees (across the world) in an attempt to mitigate climate change and in the UK to increase our percentage tree cover from a pretty low base.  Sadly, twentieth century forestry in the U.K was largely based on monocultures (for timber production). The trees planted were large stands or plantations of conifers - using Scots Pine, Larch and Spruce. These plantations not only lacked biodiversity, but were / are susceptible to wide scale pest infestation and extreme weather events.   Woodlands and forests that have a diverse range of tree species are not only healthier but show greater growth and carbon fixation. They are more resilient.  The diversity of trees ensures the each species accesses slightly different resources from the environment  - from soil minerals, water and light.  Diversity means that trees of the same species are less likely to be clustered together so pest and pathogen outbreaks are less common or less severe.  One area that has undergone an extensive and diverse planting regime is Norbury Park Estate (near Stafford).  Since 2009, over 100 different tree species have been planted, and the woodlands can now produce 1500 tonnes of new wood each year, and harvest 5000 tonnes of carbon dioxide from the air.  Not only can diverse woodlands / forests fix carbon, supply harvestable timber but they also offer areas for rest and relaxation. Whilst it is not possible to plant an 'instant' forest or woodland, it is possible to plant a range of tree and shrub species that will in time grow and mature to form a diverse and species-rich area.  As Charles Darwin said many years ago “more living beings can be supported on the same area the more they diverge in structure, habits, and constitution” [On the Origin of Species by means of Natural Selection, 1859] Managing woodlands for wildlife - see here.   N.B.  Opens a PDF.    

The Flow Country is a large area of peatland and wetland in the north of Scotland (Caithness and Sutherland). It is one of the largest expanses of blanket bog in Europe. It is characterised by deep layers of peat and bog pools.   Wetlands, like this,  are important  as They offer a significant wildlife habitat and They act as mitigating agents in terms of climate change (locking up large amounts of carbon). However, some four decades ago, parts of the Flow Country were planted with conifers,  there were financial incentives to  this planting; namely : grants were available towards the costs of planting other costs were tax deductible so investors spent only 20p for every £1 of capital invested prospectively, income from timber was to be virtually untaxed. This ‘forestation’ of wetlands is now generally regarded as a mistake, not only damaging a unique environment but leading to increased carbon emissions.  Now it seems possible that similar mistakes may be made again.    Read more...

The autumnal fall of leaves in deciduous trees is a well recognised event; their changing colours prior to being shed often make for spectacular displays - the New England Fall. Evergreens (with certain exceptions) do not undergo a similar loss of leaves but that is not to say that their leaves are forever green or permanent.  Indeed, each year, evergreens have a seasonal drop of their needle-shaped leaves, it is normal part of the tree’s cycle.  The leaves / needles of conifers have varying life spans; they are not a ‘permanent fixture’.  Many conifer needles will turn yellow then as they age, falling off the tree after one to several years. This change can be gradual or in some species quite rapid.  White Pines (Pinus strobus) typically retain their leaves for 2 to 3 years, whereas Scots Pines (Pinus sylvestris) usually retain their needles for three years. Larches, which are conifers (Larix sp) are somewhat unusual in that they shed their leaves every autumn.  The stress that a tree experiences through drought may result in more rapid browning and greater loss of leaves. Read more...

In recent times, new or different threats have emerged to upset the balance of woodland and forest ecosystems.   In the 1960’s and early 70’s concern focussed on the effects of air pollution, particularly the effects of acid rain.  This type of pollution was characterised by the deposition / assimilation of sulphur dioxide and its derivatives (sulphuric & sulphurous acid), plus various nitrogen oxides.  This air pollution was largely due to industry and traffic. Some of the most striking effects of ‘acid rain’ pollution were seen in the coniferous forests of Germany - where it was termed : Waldsterben [Wald=forest plus sterben=to die].  Read more...

The monkey puzzle tree is a popular garden tree.  It is so-called as when first seen in this country (in the mid C19th) by Charles Austin, he is alleged to have said that “it would puzzle a monkey to climb that”.  As the tree had no common or popular name, it became the ‘monkey puzzler’ or ‘monkey puzzle tree’.  Its scientific name is Araucaria araucana, and otherwise known as the Chilean Pine or Pehuén.  It is native to the Andes of Chile and Argentina.  Not only is it a long lived tree (with a life span of a thousand plus years) but the species has been around for a long time (in excess of 200 million years).  It is sometimes described as a “living fossil”. A tree can grow to a considerable height - towards 150 feet with a trunk diameter of some five feet. Read more...

The woodlands blog has previously reported on the havoc being wreaked by bark beetles.  Such beetles may be small (about half a centimetre in length) but their effects on the western forests of North America is colossal - indeed some parts have lost 90% of their conifers.   Outbreaks of these beetles have been increasing in size and severity. The beetles onward march was generally kept in check by long and cold winters, but with warmer temperatures (especially in the winter months) and a longer season for reproduction bark beetle populations have been gaining ground, even making their way into parts of the boreal forest of North America. Read more...