As stated in the previous post, bark is a mixture of living and dead cells.  Cork cells abound, especially in trees like the cork oak, where the cork may be regularly harvested.  This may be used for flooring, insulation and yes - corks for wine bottles. However, bark is a source of many things.  From early times, bark from trees like alder, buckthorn, oak, birch etc were used to make dyes for clothing.  Material from the inner bark of some trees (e.g. lime, willow, mulberry) was a source of fibres for clothing and cordage (string / yarn). Herbalists also found a use for the bark of certain trees.  Infusions of willow were used to treat fevers, the ‘ague’, rheumatic aches and pain.  It contains salicin, which the body converts to salicylic acid, an early 'form' of 'aspirin'. Interestingly, Nicholas Culpeper, in The Complete Herbal (of 1653) gives a number of uses for willow, including to staunch wounds, but does not mention pain control. The bark of the Cinchona [Jesuit’s bark] gave quinine - a treatment for malaria [caption id="attachment_39935" align="alignleft" width="300"] Amber[/caption] When a tree, like a pine, is injured (through storm damage or insect attack), the bark can produce resin - a sticky and viscous liquid.   The resin is produced in resin ducts present within the bark tissue, though the ducts may be present in deeper tissues. Research has shown that ponderosa pine trees that had more (and wider) resin ducts survived drought and bark beetle attack better.  The resin can harden and help seal wounds . Many resins contain terpenes, such as alpha-pinene and limoneme.  The resin from pine and other conifers can under special circumstances be converted into amber.  Sometimes, the resins produced can be fragrant. Trees of the genus Boswellia and Commiphora produce a aromatic resin that gives frankincense and myrrh respectively.  Both are produced by the wounding of a tree so that its resin seeps out.  Both may be used in the making of incense. Another bark exudate comes from certain species of Acacia - Gum arabic, which forms from the hardened sap (adjacent image).  Acacia species belong to the ‘Bean’ family (Fabaceae).  The gum is collected from trees, mostly in Sudan and the Sahel.  Gum arabic is a mixture of glycoproteins and polysaccharides.  The polysaccharides are constructed from the sugars arabinose and galactose. It is soluble in water and edible, and has a number of uses in the food and pharmaceutical industries. Tapping or wounding the bark of different trees can result in various fluids being released, for example, latex.    White or yellow latex is produced by the rubber tree (Hevea brasiliensis).  The latex is found in special vessels within the bark - laticifers. The process of tapping rubber trees is outlined in some detail here.   Latex production is not confined to woody trees, small herbaceous plants like dandelion and spurge can produce a white, milky latex (as can the opium poppy). [caption id="attachment_39984" align="alignleft" width="300"] Euphorbia latex[/caption] The latex produced by some members of the Spurge family can cause burning pain, inflammation or even blistering - for example that of the Pencil tree.  Such toxic saps most likely evolved to deter animals from grazing. Sometimes, a watery sap may be collected from the bark.  This is the case with Birch.  Sap may be collected (tapped) in early Spring, when sugars and other materials are being mobilised for growth, leaf production etc.  Sap may be collected later but is said to then have a bitter taste.  The sap is an interesting ‘cocktail’ of amino acids, protein, sugars (glucose & fructose), betulinic acid, proteins, vitamins C & B, and minerals.  It is used to make a much favoured drink in Northern Europe and should be consumed within days of collection.  Birch trees are quite sensitive to tapping. Not watery, but very sugary is Maple Syrup. Maple trees are tapped by drilling holes through their bark and into their trunks. Starch is stored in the trunks and roots before winter, it is then converted and mobilised in late winter / early Spring. The collected  sap (through tapping) is then heated to produce a concentrated syrup. Thanks to Montemari at Pixabay for image of gum arabic

Bark exists to protect a tree from ‘attack’ by the elements, pests, ‘predators’ (animals who would eat it) and disease causing organisms.  There is no easy definition of what constitutes bark,   a slightly technical definition might be ‘the tissues that lie outside the vascular cambium'.  The vascular cambium is a layer of dividing cells that gives rise to xylem tissue and phloem tissue.  The cells nearer the centre form the xylem, those towards the outside form the phloem.    The inner part of the bark contains various types of living cells, for example, glands that produce latex (as in natural rubber), oils and resins.  Moving outwards, there lies the rhytidome or outer bark, an amalgam of living and dead material - notably cork cells.  The cork cells fill with a waxy material - Suberin. Eventually, these cells die and form much of the bulk of the bark.  The nature of bark is immensely variable. Wind, fire and frost can seriously damage or kill trees but bark helps  to protect them.   Trees are eminently combustible as is evidenced by the recent forest fires in Australia and California. However, some trees have a very thick bark that can protect them against fire.  The cork oak has a bark that can be up to 30 cm thick, it is so thick that it can be harvested periodically without killing the trees.  Cork oak is grown extensively in the mediterranean region. Giant Redwoods too are noted for having an extremely thick bark. Their bark is very fibrous and can be up to three feet thick, it offers protection against fire (and rock fall which is also a hazard in their home habitat). In contrast to cork oak and redwoods, some trees like the eucalypts have a bark that is rich in oils and very flammable.  The bark also ‘peels’, strips are shed onto the forest floor. There are many species of Eucalyptus and several different types of bark are recognised.  [caption id="attachment_35352" align="alignleft" width="300"] Woodland recovering from a fire[/caption] If and when this oil rich bark builds up on the forest floor, it will contribute significantly to the intensity and ferocity of any fire. Indeed, it has been likened to adding petrol to a fire ’3 centimetres of leaf litter can cause a conflagration equivalent to one fuelled by a centimetre of refined gasoline’.  The leaves are also rich in oil so the crowns of the trees can also contribute to / exacerbate any fire.  The peeling or exfoliation of bark is not restricted to Eucalypts, it can be seen in trees much closer to home - such as the birch.  Its bark can be removed in long strips and has been used in covering a canoe or roofing material. Whilst bark can protect against fire, it can also deter animals - large or small from inflicting damage.  For example, there is an African species of Acacia known as knobthorn that has a bark covered with thorn-like structures.  These 'thorns' deter elephants from eating the bark.  Elephants can consume a lot of vegetation in a day and tree bark is much favoured.  A variety of animals may feed on bark material, for example deer, squirrels, and beavers, but the list could also include orang-utans, rhinos, bush babies and porcupines. North American porcupines use their large front teeth to eat bark and stems. Bushbabies generally feed on insects during the wet seasons, but during drought / dry periods - they feed on the resins / gum that flows from the trees in their woodlands. In the UK, a lot of bark damage is done by deer, especially during the winter months when other food sources are limited.  In the summer months, male deer rub their heads / antlers against the trunks of trees - inflicting damage.  Such activity can prevent regeneration in natural woodlands.  Tree guards may be needed to allow young trees to establish themselves (or fencing to create a ‘deer free’ zone).  Guards also protect against rabbit damage.  Grey squirrels can also cause damage to trees as they gnaw stems to reach the ‘sweet’, sap-filled tissues just below the bark, this activity is usually seen in late Spring and early Summer. [caption id="attachment_5312" align="alignleft" width="300"] xylem vessels[/caption] Whilst bark is broadly protective, it can also offer a home to certain pests.  Bark beetles lay their eggs below the bark so that when the larvae hatch, they can feed on the nutrient rich tissue of the cambium and phloem.  Bark beetles have been responsible for the loss of millions of trees in the United States and Canada.  The scale of the loss is much greater than in the past, when cycles of beetle infestation and fire created a mosaic across the countryside of young and old trees.  Ageing stands of trees coupled with warmer winters (which have helped the overwintering stage of the insect)  have contributed to the spread of bark beetles.  The beetles breed and feed beneath the bark, damaging the phloem and cambium tissue.  Consequently, the tree's transport systems begin to fail and the beetles may also introduce disease-causing fungi and bacteria. To a certain extent, trees are able to repair damage to their bark but the response is varied according to the nature of the damage and the tree involved. Some trees can produce ‘callus tissue’ that heals over the ‘wound’, leaving a scar. Some trees, such as the pines, produce resins and antimicrobial compounds in response to injury.  This sticky resin may trap insect invaders as is witnessed by those trapped in time capsules of amber.   Apart from bark beetles, other animals and plants live in or on bark in a variety of associations, some parasitic as is the case with fungi (like the polypores), whilst lichens and mosses are epiphytes.  They use the bark as a substrate on which to live, grabbing nutrients and water from rainwater as it trickles down.   The many uses of bark tissue can be left for another woodlands post. [caption id="attachment_39940" align="aligncenter" width="620"] Section through bark[/caption]

Bark is the term that is often applied to the outer covering of tree stems and other woody plants. It serves to protect a tree from  Water loss Insect attack Infection by bacteria and fungi Physical damage (by fire, animals, rock fall) The nature of bark is immensely variable.  In some trees, the bark is extremely rough, corrugated and thick.  In others it is is thinner and appears to peel off in strips.   Redwoods are noted for having an extremely thick bark (see featured image above). Their bark is very fibrous and can be up to three feet thick. Read more...

Spain has a lot of forestry as it is a surprisingly large country, being about twice the size of Britain and only about 10% smaller than France. The tree cover is about 30% so about twice as dense as Britain's, which means the Spanish have about four times as much forestry as we do. But it is very different in both composition and in how it is managed. For a start, 80% is owned privately and where it is state owned it is almost always owned by the local community. Read more...

Wood is such a familiar material that we tend to take it for granted.  In general, it is a long lasting, fibrous material that is found within the roots, stems and branches of trees and shrubs.  It is mainly composed of xylem – a tissue that brings water and minerals up from the roots and distributes the minerals and water to the leaves and growing tissues of the stem. Read more...