At first, you might not notice them, but ants in a woodland are important. Their presence indicates healthy woodland, and the Forestry Commission recognises them as a ‘keystone species’ in the woodland / forest ecosystem and encourages woodland owners to treat them sympathetically.  Ants, like bees and wasps, belong to a group of insects known as the Hymenoptera .  These are insects with membranous wings and narrow ‘waists’. Like bees, ants are ‘social insects’  and within their nests, there is a division of labour (workers, soldiers, queens).    There are four main species of true wood ant - Formica rufa, F. aquilonia, F. lugubris and F. pratensis. Ants are important for a number of reasons, for example :- They help disperse seeds.  They manage pests by preying on herbivorous insects that damage leaves They contribute to nutrient recycling, t Their nests provide a habitat for a whole range of invertebrates that live specifically in wood ant nests. For example, the woodlouse  (Platyarthrus hoffmannseggii), which has ‘lost’ its eyes and colour and is completely adapted to living in the dark recesses of the ants’ nests. They are a source of food for various predators including woodpeckers. Through the building of nests, they affect the soil profile, the drainage and permeability of the top soil, and help with the distribution of various nutrients.  A wood ant colony nest can persist for many years and survives so long as there is a queen / s present.   During the winter months, only queens and a limited number of the workers live on deep inside the nest. Once the temperature begins to rise in the Spring, then workers will begin to emerge and start to forage – and the queen(s) will begin to start laying eggs again.   Fertilised eggs will give rise to female ants, and unfertilized eggs give rise to males.   The males and queens have wings, and are bigger than the workers. The life expectancy of a worker is about 60 days.   Ants such as the southern wood ant [Formica rufa]. build large, ‘thatched’ nests, (see the film link below) which may contain many thousands of workers and queens (up to a quarter of a million). An ant nest is instantly recognisable, it looks like a roughly shaped dome, made from an untidy heap of leaves and twigs – with ant trails leading to and from. The heart of the nest is deep underground, whilst on the surface is the dome shaped ‘thatch’. The thatch consists of a variety materials in the thatch – small twigs, moss, heather, pine needles.   These materials are positioned so that they  not only intercept the sun’s rays to help warm the nest  but also are laid in such a way that water / rain runs away from the nest, reducing the risk of flooding.  Some ant species build their nests in /on gravel or sandy / stony soils - which reduces the risk of water logging. Like many insects, ants make use of pheromones (volatile chemicals). For example, to lay paths to the nest. The study of ant trails and ant behaviour has inspired computer programs / simulations to the “travelling salesman problem”, that is finding the shortest route round a large number of fixed locations (think of the Amazon’s driver’s day!  Woodlands TV has produced another film that focuses on woodland ants.  The film, presented by Giles Pitman, can be seen either on this website or on YouTube https://youtu.be/9wtvwlL2Wzg. Interesting fact : Ants can spray formic acid from their abdomen – when threatened. This smells like vinegar and is thought to be a deterrent to birds or other predators.  

Many organisms help to disperse seeds in woodlands or forests, from squirrels, birds, bears, foxes, rabbits.  The fruits / seeds may be eaten and pass through the gut of the animal or may be caught in their fur, or stuck to them (like the seeds of mistletoe or cleavers).  However, there is another 'army of helpers'  - ANTS.  Ants may disperse seeds in two distinct ways.  The first way is  through the activity of harvester ants.  Like squirrels and other ‘gatherers’, harvester ants move through the wood or forest gathering seeds and transporting them back to their colonies.  As they transport the seeds, some get dropped or lost on the way.  Others may be ‘cached’ / stored,  in or near the colony for later consumption, but then are ‘forgotten’.  Such seeds may later germinate and establish themselves somewhere away from the parent plant.  This accidental dispersal is sometimes referred to  as  ‘seed predation’. a second method of seed dispersal by ants involves a more intimate association between particular plants and specific ant species.Some plants, when they form the seed, produce an 'extra structure', attached to the seed; an ELAIOSOME or fat / oil body.  This elaiosome lures or attracts ants to the seed.  Wood anemone and cow-wheat (Melampyrum pratense) seeds have an elaiosome. [caption id="" align="aligncenter" width="512"] cow wheat ; thanks to Kristian Peters -- Fabelfroh 09:39, 3 October 2006 (UTC), CC BY-SA 3.0, via Wikimedia Commons[/caption] Ants need some fat, and fat-rich food is hard to come by in the woodland setting.  The ant carries off the seed to the colony, where the fat body is eaten (often by the ant larvae).   The seed is then left and can later germinate. This interaction between plant and ant is termed  MYRMECOCHORY. It is a form of mutualism, as there are benefits to this arrangement for both ‘partners’. The seeds of the plant are ‘protected’ from seed predation by birds and rodents.   The seed is dispersed from the parent plant, ants bearing a seed for some 70 metres have been recorded.  The seed is then (often) deposited in a nutrient rich microhabitat. The ant gains a ‘tailor made’ food material rich in fat, an energy rich material.  In some areas, such as Australia, the seed may be protected from fire as it is buried at some depth. Myrmecochory is seen in many species in north eastern America, but also Europe, Australia and Africa. wood ants https://www.youtube.com/watch?v=ZVGcl4Om-C8&t=40s    

Sadly, the world is losing species, both plant and animal, at a significant rate.  Indeed, some claim that we are now experiencing the sixth mass extinction.  In contrast to previous extinctions (the Permian extinction is thought to be due to an asteroid impact), the present loss of species is largely associated with a mix of direct and indirect human activities. These include :- destruction and fragmentation of habitats,  Exploitation fishing stocks and hunting (think dodo),  chemical pollution,  invasive / introduced species, and  human-caused global warming The loss of animal species has knock-on effects in terms of food chains and biodiversity. Plants are also affected as many rely on animals for the dispersal of their fruits and seeds.  In times of global warming, it is essential that plants can reach new areas that are suitable for their growth.  If not, they are stuck in areas where they may not be able to survive in the changed / changing conditions.  This could mean that plant species are lost, together with the ‘ecosystem services’ that they provide (be it food, timber, carbon storage, flood mitigation etc).  Seed dispersal is also important in terms of recovery from ecological disasters, like wildfires.  Natural forest regrowth usually happens through seed dispersal. If an ecosystem is rich in species, it is generally more resilient to environmental change.  The relationship between fruit / seed dispersal and animals has been significantly affected by the creation of roads, motorways, farms, and the development of cities - essentially habitat fragmentation.  Animal dispersal is often associated with fleshy fruits.  Whilst this is particularly true / obvious for many tropical fruits, it is also the case for many plants in temperate regions.  Berries, hips and haws are dispersed by animal means, with birds being particularly important agents. Several pines produce large seeds and attract corvids such as nutcrackers and jays. The birds, sometimes called scatter hoarders, collect seeds and bury them in areas away from the parent trees but in habitats suitable for the next generation of trees  Mammals also play significant roles.  In Africa, elephants are important  seed  dispersers  for  numerous  species; they  have an extra-ordinary sense of smell and will search out ripe, fleshy fruits. Some seeds have been shown to be distributed 60+ kilometres from a parent plant. Not only this, but the journey through the gut of the elephant seemingly increases the chance of germination, and being deposited in the dung reduces the chance of the seed being eaten by beetles.  Some monkeys in South and Central America eat as many as fifty different types of fruit in a day. carrying some off in their stomachs and dropping others to the ground. In Britain, as part of their diet, foxes will eat various wild fruits, like blackberries; squirrels eat nuts; and mice / voles eat grass and other seeds.  Even invertebrates, like ants, disperse seeds. This may be through the activity of harvester ants, which, like squirrels and other ‘gatherers’,  forage the ground of the wood or forest (collectively) gathering large quantities of seeds and  then transport them back to their nests / colonies.  As they transport the seeds, some get dropped or lost on the way.  Others may be ‘cached’ in or near the nest for later consumption but then are ‘forgotten’ or ignored. Some fruits contain seeds covered with a sticky substance as is the case of Mistletoe. When birds feed on the fruits, the seeds often stick to the beaks of birds.  Then, they may wipe the sticky seed off on a branch;  or it may be eaten and pass out in the bird’s droppings. The ‘glue’ (viscin) around the seed helps fasten the seed in place. Even humans carry seeds far away for plants, for example, by taking an apple on a picnic and throwing the core with its seeds into the bushes. Or seeds may transported in the mud sticking to boots and shoes, or indeed on tractors, cars or other machinery. The loss / extinction of animal species from any given habitat will sooner or later effect the plants.  We are only beginning to fully appreciate the interdependence of life.    The loss of any species - plant or animal - will undoubtedly have unintended and unforeseen consequences which can only be to the detriment of all life on earth.  

In general, as plants are fixed in place, they have a bit of a problem when it comes to reproduction. That is, exchanging genetic material with one another and then ensuring that their offspring can establish themselves away from the parents so that they are not in direct competition.   Some plants have solved this by using the wind to move pollen and/or seeds (anemochory), some use water (hydrochory) and many others have ‘entered into associations’ with animals.    The list of animals providing ‘assistance’ is quite varied.  In the case of pollen, transfer is achieved by beetles, bees, bumblebees, hoverflies, hummingbirds, fruit bats, moths etc.  The dispersal of seeds by animals is known as zoochory.   Read more...

That seeds are dispersed by animals is well known; most elementary texts on biology are littered with a variety of examples. Fruits (and their seeds) attract animals with the edible parts; from fleshy fruits - plums, mangoes, berries,  cherries etc for frugivores, to various nuts.  As the animal searches for and collects the fruits (monkeys in the forests, squirrels in the woodlands), some may be lost or discarded on the journey ‘home’.  Other fruits / seeds cling to an animals by attaching to their fur / clothing using various hooks or barbs – a classic example being cleavers or goosegrass. Some seeds are dispersed incidentally; they may be lurking in muddy soil and adhere to the hoof or foot of an animal, bird or human, or be carried some distance by car or tractor tyres. However, many species of plants have an ‘unseen’ army of helpers – ANTS.  Ants disperse seeds in two distinct ways.  Read more...