We can communicate in many ways, other animals may  message each by vocalising or gesturing.  Some release air-borne chemicals (pheromones) to attract a mate.  Plants also communicate by chemical signals that they release into the environment. The most obvious example is the release of scents to attract pollinators.   These floral scents are usually oils, made from volatile organic compounds (VOCs).  VOCs are carbon based compounds, such as the terpenes. They usually have a strong and pleasant odour.  They form partt of the scent of many plants, for example, honeysuckle, roses, jasmine, lilac, and mock orange.  Different plants produce different scents.  These different chemical signatures help pollinators identify particular species, and locate the nectar on offer. The VOCs produced by ripe fruit help attract animals that assist in seed dispersal. [caption id="attachment_43370" align="aligncenter" width="675"] Buff tailed bumblebee visiting a foxglove[/caption] VOCs are also produced by leaves when plants are under attack.  These may deter the attacking organism directly, e.g. caterpillars, or they may attract an organism that predates upon the herbivore. When pollen beetles feed on oil seed rape, the plants release VOCs which attract the attention of insects (a type of wasp). The wasps lay their eggs in the larvae of the pollen beetles. The pollen beetle larvae are then ‘eaten alive’, by the developing wasp larvae.  Sometimes, VOCs are produced by stressed or attacked plants,  these VOCs ‘warn’ nearby plants of the same species. These plants can then activate their defences before they are attacked. This helps ensure the survival of some of the population. Recently, it has been suggested that mycorrhizal systems act as a physical and chemical ‘means of communication’ between the trees in a woodland, in what has been termed the ‘wood wide web’. The term was coined by Dr. Suzanne Simard, a forest ecologist at the University of British Columbia to describe the complex relationships between fungi and plants in woodland ecosystems.  Interestingly, VOCs can influence air quality and atmospheric processes.  In a woodland,  the scent of pine or other conifers may permeate the air.  Pines release volatile compounds, such as alpha-pinene.  Recent research has established that these volatile compounds / vapours are not only responsible for the characteristic scent, but also contribute to the formation of aerosols in the atmosphere* both in and around such woodlands and forests.  In the presence of sunlight, VOCs can interact with the gas nitric oxide to form ground level ozone, which contributes to smog formation. Forests can be a significant source of VOC emissions. The production of VOCs is affected by environmental  conditions.  VOCs release generally goes up with an increase in temperature, as the plants metabolism increase.  An increase in temperature also helps in the evaporation / volatilisation of plant oils and scents. Water availability and light intensity also affect VOC release.  * An aerosol is a ‘mixture’ of very small particles (solid or liquid) in air; other examples of aerosols include mist, cigarette smoke, or car exhaust fumes Addendum : Heavier and far rarer VOCs are the sesquiterpenes, larger molecules that trees in the Amazon release when exposed to drought.  These reactive molecules help form tiny particles that seed clouds and scatter sunlight. These emissions could change the weather and climate over the whole region.  For full details see : https://www.earth.com/news/amazon-trees-emitted-never-before-seen-chemicals-during-drought/  

Many plants have a distinctive scent, think of sweet peas, jasmine or honeysuckle, or stand next to a pine tree on a warm, summer’s day.  The scent is due to the release of volatile organic compounds (VOC’s, often oils), produced by specific tissues or glands.  Often it is the nectaries within flowers that produce the scent, apart from their ‘job’ of producing the sugary nectar.  The nectaries may be found on almost any structure within a flower - petals, sepals, stamens, ovary*. The location of nectaries varies from species to species.  There are other structures that can produce scent, for example, trichomes, and osmophores. Osmophores are clusters of cells specialising in scent production.  Any part of a plant can release scent, for example, the leaves of eucalyptus, lavender or myrtle. The scent of a plant may include a variety of VOC’s, indeed there may be dozens of different organic compounds contributing to a particular scent.  Many of these compounds are terpenoids (isoprenoids).  They contribute to the scent of eucalyptus oil, lavender oil and the flavours of cinnamon and ginger. Scent may have a number of functions.  It may be released to attract specific pollinators - moths, butterflies, bees, hoverflies etc. (who have learned to recognise the scent).  The production of VOCs can be modulated, for example,  scent production may be turned off when a flower is pollinated.  A scent may also unfortunately be a signal to herbivorous insects to ‘come and feed’. So, scent have positive or negative effects. A scent may be produced to deter herbivory by certain insects.  Sometimes, plants have a different approach. For example,  when pollen beetles feed on oil seed rape, the rapeseed releases VOCs which attract the attention of other insects.  Specifically, those that will lay their eggs in the larvae of the pollen beetles. These insects are usually from the same family as bees, wasps and ants - the Hymenoptera (insects with membranous wings and a ‘narrow waist’).   The pollen beetle larvae are then ‘eaten’ from the inside by the developing parasitoid larva.  The release of VOC’s is affected by a number of factors temperature, light, circadian rhythms, physical damage and drought.  As the temperature increases so the amount of VOCs released increases (usually). This may be experienced in coniferous woodland.  Conifers give off a variety of volatile oils (i.e. biogenic VOC’s) that contribute to a unique aroma and the formation of aerosols found in the air in and around such woodlands and forests; it is most noticeable in warm weather.  [An aerosol is a ‘mixture’ of very small particles (solid or liquid) in air; other examples of aerosols include mist, cigarette smoke, or car exhaust fumes]. In snapdragons, the most scent is emitted at noon which tends to coincide with pollinator activity, in contrast tobacco plants scent release is in the evening / night when hawkmoth are active.  Drought reduces the ability of plants (like rosemary and thyme) to produce / release VOC’s, this in turn, has been observed to affect which pollinators visit their flowers.  Nectaries located within the flowers of a plant are sometimes referred to as nuptial nectaries, whereas those found in other parts are termed extra-nuptial.